The extended phenotype is a sequel to the selfish gene. The extended phenotype 1982 offers an alternative view on biology and the process of evolution. Among his previous books are the ancestors tale, the selfish gene, the blind watchmaker, climbing mount improbable, unweaving the rainbow, and a. The long reach of the gene, richard dawkins, oxford university press, 1989, 307p, trade pb, covers bumpedscuffedcreased, clean text, solid binding8. Then the extended phenotype displaces the individual organism as the locus of expression of genetic material. I want to argue in favour of a particular way of looking at animals and plants, and a particular way of wondering. Manipulation of host behavior is such an adaptation. The extended phenotype is one of the biggest ideas in evolutionary biology and examples are all around us. Along with the illuminating insights of the selfish gene and memes, dawkins gives us the extended phenotype.
The main topic tackled in this book is the level at which natural selection acts. When we say that a gene is a gene for something, all we mean is that possession of that gene as opposed to its competitor for that locus makes a difference regarding that something. The extended phenotype the long reach of the gene by richard dawkins with a new afterword by daniel dennett oxford university press, 1999 not your parents web of life in the 19th century, thomas. It is a fascinating look at the evolution of life and natural selection. Richard dawkins the extended phenotype the long reach of. Extended phenotype hopi hoekstra harvardhhmi youtube. His bestselling books include the selfish gene 1976, the extended phenotype 1982 and its sequel the blind watchmaker 1986. The extended phenotype by richard dawkins waterstones. Dawkins first book, the selfish gene, was a smash hit. By the best selling author of the selfish genethis entertaining and thoughtprovoking book is an excellent illustration of why the study of evolution is in such an exciting ferment these. The extended phenotype is a 1982 book by the evolutionary biologist richard dawkins, in which the author introduced a biological concept of the same name. Breaking with the darwinian paradigm that puts the individual organism center stage, author richard.
In the selfish gene, richard dawkins crystallized the genes eye view of evolution developed by w. The selfish gene is a classic exposition of evolutionary thought. The expression of an adaptive extended phenotype sandra b. The order of emergence brand, complex systems, culture, extended phenotype, gene, language, meme, meme complex, phenotype, public relations 2 life posted on 16. The idea of the extended phenotype ep, which was first proposed by the british evolutionary biologist richard dawkins to explain how and why organismsor, more fundamentally, their genesare able to. In the extended phenotype, richard dawkins proposes that the expression of a gene is not limited simply to the organisms physical appearance or phenotype, that is the direct synthesis of proteins, or to the organisms behaviour, but also includes the impact of the phenotype and the behaviour on the organisms environment. An extended phenotype is the behavioural phenotype of a gene optimon.
One example is burrowing behaviour in oldfield mice, on which a genetic study was. The term covers the organisms morphology or physical form and. The writer of the extended phenotype is an english biologist. Dead ants were found under leaves, attached by their mandibles, on the northern side of saplings. We saw that to talk of an individual behaving so as to maximize its inclusive fitness is equivalent to talking of the gene or genes for that. The extended phenotype is a more difficult read for the nonscientist that the selfish gene but it is manageable with some rereading of key passages. This view of the gene has an interesting analogy in archaeology. In the extended phenotype, richard dawkins proposes that the expression of a gene is not limited simply to the organisms physical appearance or phenotype, that is the direct synthesis of proteins, or to the.
Extending the extended phenotype article pdf available in biology and philosophy 193. We studied the fungus ophiocordyceps unilateralis, a locally. Phenotype from greek pheno, meaning showing, and type, meaning type is the term used in genetics for the composite observable characteristics or traits of an organism. Science the extended phenotype is a sequel to the selfish gene. He was the first holder of the charles simonyi chair of the public understanding of science at oxford, a position he held from 1995 until 2008, and is emeritus fellow of new college, oxford. The extended phenotype carries on from where the selfish gene takes off. Specialized parasites are expected to express complex adaptations to their hosts. An extended phenotype might be understood as any effect of genes extending beyond an organism into the world around it which also provide fitness for their replication. Pphenotype is the set of observable characteristics of an individual resulting from the interaction of its genotype with the environment. All effects of a gene upon the world where the effects influence the survival chance of a gene richard dawkins. Chapter 10 discusses various notions of individual fitness, and concludes that they are confusing, and probably dispensable.
The extended phenotype by richard dawkins october 14, 2008 at 19. Why the evidence of evolution reveals a richard dawkins. The extended phenotype how richard dawkins got it wrong. I dont usually like multiple narrators maybe because of librivox with a different narrator per chapter, but for this book it works well as each character reads their own letters. Adaptation and the natural selection that follows from it depends upon a progressively harmonious. Hoover k1, grove m, gardner m, hughes dp, mcneil j, slavicek j. Clinton richard dawkins is not just a writer but an ethologist and a very good evolutionary biologist as well. According to wikipedia the book is set in 1946 and is an epistolary novel, composed of letters written from one character to another.
The first thing i will say about the extended phenotype ep is that it is far from the first book on evolution you should read and as a standalone dawkins book it is a poor choice because it is. But dawkins clear exposition is accessible to all who are prepared to put in a. Dawkins believes it is important because, of all the factors and influences involved, the gene is the only one that persists, that survives the death of the organism. The extended phenotype the long reach of the gene is the book richard dawkins wants you to read if you read nothing else of mine. In genetics, the phenotype of an organism is the composite of the organisms observable characteristics or traits. The extended phenotype richard dawkins audiobook youtube. An extended phenotype is an animal artefact, such as a nest or burrow, that is influenced by a gene and is a measure of behavior. This hypothesis is not experimental in the traditional sense. Audiobook publishing made easy alexa actionable analytics for the. A reply to laland, turner and jablonka richard dawkins university museum of natural history, university of oxford, uk i am grateful to the three. Dawkinss theory is that individual organisms are replicators that. The extended phenotype is a conceptually deep book that forms important reading for biologists and students. The extended phenotype was written to reinforce the view that the gene is the most important element in evolution.
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